Mark was born to W. Morgan Sheppard and his wife on the 30th of May, 1964 in England. He is also known for being doubled casted with his father as they play the younger and older versions of the same character on several movies.
Born in London, England, Mark is of an Irish-German background. At the age of 15, he was interested in music and became a professional musician. He toured with bands like Television personalities and Light a Big Fire and reacted several songs. After moving into the United States, Sheppard auditioned for the production of Cock and Bull Story and won the 1992 L.A. Drama Critics Circle Award as well as the LA Weekly and Dramalogue Awards for his role there.
For television, he has starred in The X-Files, Soldier of Fortune and has had recurring roles in shows like The Invisible Man, The Practice, Special unit 2, The Chronicle, JAG, Star Trek: Voyager, Monk, White Collar, CSI, Leverage, and Chuck among many others. The TV series Charmed has starred him as well. His other appearances include acting as an FBI agent on Joss Whedon's show Dollhouse, and, ironically, a notorious criminal on Whedon's show Firefly. He has also appeared as a villain on 24 and on Medium. His other television appearances include acting in The Middleman, Bionic Woman, Doctor Who and Battlestar Galactica. He started acting as demon Crowley on Supernatural since 2005 and has been doing so until now. His film credits include acting for In the Name of the Father, Unstoppable, Broken, New Alcatraz and Megalodon. He has also directed Mysterious Island.
As far as his personal life is concerned, Mark is currently married to Sarah Louise Fudge. Previously, he had been married to Jessica. Mark and Jessica married on the 6th of March, 2004. After having two children named Max and William together, the couple had marital problems and thus got divorced in 2014. Following his divorce, he started a relationship with Sarah Louise Fudge. Sarah, an Australian mining heiress, met him at a Supernatural convention. Mark and Sarah became engaged on August 20, 2015. They announced that they were expecting their first child on September 26, 2015. Following this announcement, the couple got married on 9th November, 2015 in Denver, Colorado. Their baby girl, Isabella Rose Sheppard, was born on March 1, 2016.
He became a United States citizen in 2015.
Mark is sometimes credited as Mark A. Sheppard.
- In response to a fan's question, Mark said he was a "Dean-girl".
- Mark coincidentally shares his birthday with co-star Jensen Ackles' daughter, J.J.
- Mark's son Max Sheppard made an uncredited appearance as a demon in season 10 episode "Soul Survivor".
- Mark expressed displeasure over Crowley's death in Season 12, believing he deserved better.
- Mark's second son William was born on June 16th (he's 16 as of 2021). Mark refers to him as "mysterious".
- Coincidentally Mark already had portrayed a demon before. It was the demon Arnon in TV series Charmed.